Born 1950 in Oldenburg. Study of biology and mathematics in Bielefeld and Bochum (1978), German Habilitation in zoology in Bochum (1985), Heisenberg Scholarship Oxford Univ./UK (1986-1987), Professor of Zoology and Behavioural Ecology Bern Univ. (1987), Director and Scientific Member at the Max Planck Institute of Limnology (1999-2007), 2007 to 2018 Max Planck Institute for Evolutionary Biology. Honorarprofessor of Kiel University (since 2000). Emeritus in March 2018.