Born 1952 in Salzburg, Austria. Study of biology, doctorate (1982), postdoc in biology MIT, Cambridge, Mass. USA (1982-1986), Assistant Professor Univ. California, San Francisco, USA (1986-1992), Associate Professor HHMI, San Francisco, USA (1992-1995), Professor and Vice Chairman HHMI, San Francisco, USA (1995-1999), Professor and Director Gene Center Munich (1999-2004), Director and Scientific Member at the Max Planck Institute of Immunobiology and Epigenetics, Freiburg (since 2004). Emeritus Director since 2021.